The shrinking and separation of the cytoplasm from the cell wall due to exosmosis of water from the protoplast.
Adapted from: Tronsmo, A.M., Collinge, D.B., Djurle, A., Munk, L., Yuen, J. and. Tronsmo, A. 2020 Plant pathology and plant diseases. CABI, Wallingford.
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